Assistant in Biology 1895-1896; Instructor in Biology 1896-1903; Assistant Professor in Industrial Biology 1903-1909; Associate Professor in Industrial Biology 1909-1912, in Industrial Microbiology 1912-1914; Professor of Industrial Biology and Microbiology 1914-1942; Acting Head Department of Biology and Public Health; 1921-1922; In charge of the Department 1922-1942; Dean of Science 1932-1942; Honorary Lecturer in Biology 1942-1944;Professor Emeritus 1944-1962.